The Hub was the seat of power of the Cybertronian Empire. A powerful planet-spanning network of cyberformed structures, it housed the Decepticon progenitor known as the Liege Maximo. Inhabiting it were a wide variety of mechanical organisms, beings that might have once been Decepticons, but had been advanced to the point of having multiple forms and reduced to a hive mentality, little more than extensions of the Liege Maximo's will.
The Hub had long been performing a complex operation through successive iterations, its accuracy improving as planets were added to the network. The goal of this operation was to allow the Liege Maximo to ascend to the realm of the Dark Gods. At a critical moment this operation was disrupted on two fronts by a small group of Autobots led by Grimlock and an alliance of Decepticons and Autobots led by Ultra Magnus and a resurrected Megatron. Megatron actually managed to wound his ancestor before perishing, which caused the Liege Maximo to panic and attempt to enter the portal prematurely. It collapsed, destroying him. The resolution of this threat led to the signing of the Pax Cybertronia and the end of the Great War.