Optimus Prime (Tyran)/208.28 Gamma

Optimus Prime is a character from the Transformers comics.

History
When war broke out, Optimus studied the entire history of intergalactic warfare from any datatrack he could find - he studied whole campaigns to desperate last stands, gaining a focused "playbook" of warfare strategy. He understands warfare on a practical and psychological level, and in desperate battles knows to show no mercy to opponents beyond a swift death.

Optimus was the focus of constant attacks by assassin drones and killed them all. In the aftermath of one, he witnessed a prophetic vision of the Mission City battle and realized the All Spark's ejection could spread war to other planets—but too late to stop the launch...

Almost immediately after the final battle against Megatron, Optimus Prime had to lead the Autobots against a horde of zombie Decepticons. Despite having no idea how they could be stopped and having had his power drained by fighting Megatron, Optimus continued to fight the monsters rather than let them run amok. Ratchet worked out a solution just in time before Optimus fell.

In the alternate reality of Twilight's Last Gleaming, Megatron defeated Optimus and took the All Spark at Mission City. Prime's body was now partially frozen and in Decepticon hands. He was the last and only hope of the Earth resistance, and so Bumblebee and Mikaela Banes were not happy to find him offline. She thawed him out while Bumblebee distracted Megatron, and the revived Prime challenged Megatron to a solo duel... Unfortunately, Megatron's link to the All Spark meant Optimus could not do any lasting damage and was never going to win. Luckily, Bumblebee and Mikaela saved the day, hitting the distracted Megatron with an old Sector Seven nanovirus. (Which means they were the last and only hope, but oh well.) Despite Megatron dying as a result and the All Spark being destroyed elsewhere, Prime could take no satisfaction in victory: this battle was won, but the war remained...

He became resigned to the fact that Humanity, battered by Decepticon occupation, would view both Autobot and Decepticon as hostile invaders. When NATO was manipulated into attacking the Autobots, Optimus refused to allow his men to return fire, and after the crisis passed, he was sure that the Decepticons' unknown new commander was up to something bigger. Optimus has also placed the last All Spark fragment in the care of Ratchet, though he's worried exposure to Megatron may have corrupted it.